Nell Gunn, Snowball and Esther will be back, along with more characters from Britain's biggest beauty factory, as they battle the twists and turns of war in Stratford. Will Nell find lasting love? Will Snowball reveal what forced him onto the streets? And will Esther be able to tame the wild child she takes under her wing as part of her new bursary? As flowers and vegetables blossom in the Shoot's new Dig for Victory allotment, can the inhabitants really bury their pasts?

Bristol, 1939. School leaver Maisie Miles suspects her father, a small-time crook, has an ulterior motive for insisting she gets a job at the W. D. & H. O. Wills tobacco factory but keeps it to herself. She's befriended by effervescent Phyllis Mason and kind-hearted Bridget Milligan who take pity on her and take Maisie under their wing. But beneath their happy-go-lucky exteriors, they all harbour dreams and worries about what the future holds. 

1942. Sixteen-year-old Poppy Percival turns up at Trout's clothing factory to start work as a seamstress. Poppy harbours a secret - one from which she is still suffering. But she's not the only one with a secret. Each of her new friends at the factory is hiding something painful. The war will throw their lives into turmoil but also bring these women closer to each other than they could ever have imagined.

Annie, Rose, Pearl and Millie carry on making matches for the British Army, with bombs raining down around them. Inspired by the Dig for Victory campaign, Annie persuades the owners to start Bryant & May allotment in the factory grounds. With plenty of sweat and toil, the girls eventually carve out a corner of the yard into a green plot full of life and colour. In the darkest of times, the girls find their allotment a tranquil, happy escape. 

In 1940s London, each cobbled street is strewn with ghosts of soldiers past. And for the women, their battles are on the home front. Flossy is embraced by the colourful mix of characters working at Trout's, who have turned their sewing expertise to war work. Dolly Doolaney, darling of the East End, helps Flossy settle into wartime life. All of the sewing bees play their part in defending the front line, but as the Blitz hits London, can Dolly manage to contain the secret that binds them all?

It’s 1941, and as the Second World War rages on, girls from all over the country are signing up to the Women’s Land Army. Renowned for their camaraderie and spirit, it is these brave women who step in to take on the gruelling farm work from the men conscripted into the armed forces. When Yorkshire mill girl Una joins the cause, she wonders how she’ll adapt to country life. Luckily she’s quickly befriended by more experienced Land Girls Brenda and Grace. 

Maureen Jackson is a prisoner of her father's blackmail. Three years ago, she'd been hoping to marry Rory, the man of her dreams. However, after her mother's death, Maureen was left to care for her overbearing father. Now Rory is back in London with his pregnant wife and reminds Maureen of the life that should have been hers. With the war looming, Janet Ashley hopes to marry her sweetheart Mike, but her father refuses to grant them his blessing. Janet finds herself pregnant and her mother Peggy is determined to hold her family together.

It is 1939, and working-class Evie Bishop has received a scholarship to study mathematics at Oxford when tragedy turns her life upside down. Evie must seek a new future for herself and, inspired to contribute to the war effort, joins the Women’s Auxiliary Air Force as an Ops Room plotter. Posted to a fighter station on the Sussex Coast, Evie befriends two other WAAFs. But the German bombers seem to know more than they should about the base’s operations, and soon Evie, May and Jess are caught up in a world more dangerous than they ever imagined. 

December 1930. With the Depression biting deeper, it looks like many children in the East End will wake up to nothing on Christmas morning. Widower Robbie Graham earns just enough to put food on the table for his children, Ben and Ruthie. Local cake shop owner Flo Hawkins can't bear the thought of any child having nothing for Christmas. Along with her sister, Honour, she hatches a plan to bring some festive cheer to the community.

The war is almost over - and it's up to the East End Angels to keep the home fires burning! Frankie's fiance, a doctor, is away looking after the troops in Europe - will he return safely home? Winnie has a happy secret - but can she carry on at Station 75 when she's going to have a baby? Bella is intrigued by her new friend, a Polish airman. As the war ends and victory is in sight, what's next for the girls of Station 75?  

In February, 1922, at the western-most entrance to Victoria Station in Manchester, a massive plaque was unveiled. Beneath a vast tiled map showing the lines of the Lancashire and Yorkshire Railway network, a series of seven bronze panels recorded the names of the men of the Lancashire and Yorkshire Railway who gave their lives for King and Country in the Great War - a total of 1,460 names. In March, 1940, a group of women of varying ages and backgrounds stands in front of the memorial, ready to do its bit in this new World War.... 

Sunderland, 1940, and the women go to the shipyards to do their bit for the war effort. Polly never dreamed she would be able to work in the shipyards like the men in her family but times are tough and her new job ends up giving her more than she ever expected when she meets enigmatic dock diver Tommy Watts. During the day, head welder Rosie teaches her fledgling flock of trainees their new trade, but at night she hides a secret life. And mother hen Gloria signs up to escape her brutal husband, but finds she cannot run....

Newly married to dashing RAF officer Tom, Irene Farley leaves behind her safe countryside life to move in with his family by the docks in South Shields. Little prepares her for the devastation the Jerry bombers have wreaked on the Sixteen Streets or that they would be living under her mother-in-law's roof, alongside Tom's three brothers and two wives!

Two very different girls from this tight-knit community join up as ARP Wardens to do their bit for the home front. Violet Simms lives with her controlling, widowed mother who runs the local pawn shop. At just 20-years-old, Violet longs for friendship, love and escape. It seems her dreams might come true until tragedy strikes on one of the very worst nights of the Birmingham Blitz. Grace Templeton is the eldest in her family of 10 children. Spirited Grace is determined never to become burdened by child bearing and drudgery like her mother. 

Liverpool, 1939. Bella Rogers thought her life was just starting; she can’t wait to get married to her childhood sweetheart, Bobby. But when war is declared and Bobby is called up, his letters stop, leaving Bella heartbroken and alone. Then her family is torn apart by tragedy when her angelic 5-year-old sister dies from a terrible illness. They can barely afford a funeral, but the church is packed with the entire community, devastated for their loss. Grieving for her sister and lost love, Bella finds a job at the Bryant and May factory, making matches.