Distant rumblings of conflict in Europe have reached even the secluded, snow-dusted mountains of Virginia, where Lauren Greenwood faces a battle of her own. The Great Depression is crippling America, leaving millions of its victims without shoes on their feet and clothes on their backs. Hope House - the orphanage Lauren runs - is suffering more than ever. The one thing Lauren is not short of is love. But with just a handful of dollars to her name, every day is a struggle to feed the orphans and keep a roof over their heads. Yet she refuses to give up. 

Bridget Collins is in dire straits - she needs to get out of New York, fast. With two young siblings under her wing, her options are limited. Her priest sends her as an outplacement agent on the orphan trains that run from New York to out west. With almost 40 orphans under her care, she's relieved that fellow and more experienced outplacement agent Carl Watson is there to guide her. But Carl is dealing with his own trauma and finds it difficult to handle the pain the orphans are dealing with.

Aurora Pettigrew has a loving family, a nice home, a comfortable life. She's waiting for the right man to offer marriage, and the man for her is Reid Sinclair, heir to the Sinclair fortune. But Reid's mother is against the match and unearths a secret that will tear Aurora's world apart. Unwilling to bring shame on her family and needing answers, Aurora moves to York. By chance, she reconnects with a man from her past, and before he leaves with the army to war in South Africa, he offers her security through marriage.

Chase Logan lives by no man’s leave, a solitary drifter who closed off his heart long ago - until the day he impulsively does a kindness for a beautiful young widow, and everything changes. For two years, Jessie Strong has dreamt of the day she could bring little Sarah home from the orphanage and call the child her own. But when her husband dies, Jessie fears the family she’s longed for is slipping away - until Chase Logan unexpectedly comes to her rescue. By pretending to be her husband, Chase can make all of her dreams come true.

Yorkshire Dales, 1850. As a terrible storm rages, Annabelle Wallis is shocked to find a distressed young woman at her cottage door, heavy with child. Moments later a baby girl is born. But by dawn, the mother has vanished, leaving behind the helpless child wrapped only in a silk peacock shawl. When news spreads that Lady Eliza Hartley, sister to a wealthy estate owner, has been found dead, Annabelle realises the terrible secret she has stumbled on. Terrified she’ll be blamed for Eliza’s death, Annabelle flees to the slums of York, where she plans to raise the orphan as her own.

After escaping the grip of the workhouse, Lily has kept her fiancé's business afloat while he is away fighting on the Western Front. Still battling on, she's now doing her bit for her country as an auxiliary nurse - but one thing above all else continues to weigh heavily on her heart: her long-lost sister. Born just before her mother died, the scandal was hushed-up and the baby spirited away. But now, at last, there is hope Lily could find her little sister for she has a clue to go on: the name of the notorious baby farmer who bought the child all those years ago. Mrs Jolley.

Danny gets a terrible start in life and runs away from his drunken father when he thrashes him to within an inch of his life. But life on the streets is no better, and he falls into the clutches of evil men, finding that being big and strong for his age is no protection from the people who want to do him harm. Help comes in the form of Constable Jones, who is determined to help out the children who need it, and new friends Ron and Cassie, who are also runaways. Can Danny’s dream of a loving home ever be a reality?

Clara Newton is the new Housemother of Shilling Grange Orphanage. Many of the children have been bombed out of their homes and left without families, their lives torn apart by the war, just like Clara’s. Devastated by the loss of her fiancé, a brave American pilot, she is just looking for a place to start again. But the orphans are in desperate need of her help.

When Ethan Postlethwaite, his wife, Grace, and their daughter, Amy, announce that they will be leaving the family home in the Yorkshire Dales, Grace’s parents are heartbroken. Hoping for a new life prospecting for gold in the wilds of Canada, the young family say goodbye and set sail across the Atlantic in search of a brighter future. The journey there proves hard and treacherous; upon arrival, it becomes apparent that the riches they had been promised in the gold fields have already been plundered.

London, 1910. Lily is 10 years old when she realises her grandmother, a washerwoman, is seriously ill. She's determined to do what she can to help and keep her grandmother's illness a secret - even from her beloved twin, Artie. When tragedy strikes and the twins are faced with the prospect of a workhouse or an orphanage, a benefactor offers to take Artie in and educate him. All Artie's needs will be taken care of - but the gentleman has no use for a girl. As the orphan twins grow up and take different paths, their new lives are beyond anything they could have imagined. 

As a young girl, Lillian Walsh lost both her parents and a younger sister. Now in her 20s, after enduring the death of her adoptive mother, Lillian must find her place in the world. Just as her adoptive father is leaving for an extended trip to his native Wales, a lawyer appears at the door to inform Lillian that she has inherited a small estate from her birth parents - and that the sister she had long believed dead is likely alive. 

1904: Ellen Copley is still a child when she leaves behind the sooty rail yards of Glasgow and crosses the Atlantic Ocean with a heart full of dreams. Yet within weeks of their arrival in America, her father has disappeared - leaving Ellen with resentful relatives, feeling alone and scared for her future. But then her kind Aunt Rose invites Ellen to stay with her large family in their rambling house on beautiful Amherst Island, which nestles like a jewel in the blue waters of Lake Ontario.

Left suddenly penniless, the Honorable Sophia Grafton, a viscount's orphaned daughter, sails to the New World to claim the only property left to her name: a tobacco plantation in the remote wilds of colonial Virginia. Enlisting the reluctant assistance of a handsome young French spy - at gunpoint - she gathers an unlikely group of escaped slaves and indentured servants, each seeking their own safe haven in the untamed New World.

In early 20th-century Belfast, working class Meg Preston struggles to accept her own sexuality and yearns for forbidden love. Battling the customs and hardships of their time, Meg pursues a relationship with her childhood friend Lillian Watson. But soon, tribulations of war, violence, and emigration threaten to tear everything apart.

1941. The Blitz rages over London. And even in Cornwall, the war is being fought. When Violet loses her sister in the Blitz, she must take her nieces to safety in Cornwall. On the coast, she meets carefree chorus girl Eva, who is also running from the dangers of London. But Porthcurno hides a secret military base, and soon Violet and Eva realise there’s a battle to fight in Cornwall, too. Together with local Hazel, who works on the base, they must come together to help the war effort. But will their friendship be enough to keep them safe?