Summer 1918. After their experiences on the front line of battle, young couple Amy and Edmond Derwent have now settled back in England. The Great War rages on, and Edmond, still struggling with the effects of the injury he sustained at Ypres, feels the guilt of remaining at home. But life at Larchbury is not without its own problems - food becomes scarce, and the Spanish Influenza causes deaths throughout England. Can Amy and Edmond keep their love strong? 

Two very different girls from this tight-knit community join up as ARP Wardens to do their bit for the home front. Violet Simms lives with her controlling, widowed mother who runs the local pawn shop. At just 20-years-old, Violet longs for friendship, love and escape. It seems her dreams might come true until tragedy strikes on one of the very worst nights of the Birmingham Blitz. Grace Templeton is the eldest in her family of 10 children. Spirited Grace is determined never to become burdened by child bearing and drudgery like her mother. 

Laura Cannock is on the run. Suspected of killing her bullying husband, his family are on a merciless prowl for revenge. Fleeing from her beloved home of Bolton to Manchester, Laura seeks refuge with her coal merchant uncle. But it soon becomes clear that a roof over her head comes with a price - of the type so unbearable she must escape once more. Destitute and penniless, a stench-ridden housing court in the back streets of the factories is Laura’s only hope of a dwelling - a place where both the filth and the kindness of neighbours overwhelm.

Liverpool, 1917. When Irish seaman Michael Gallagher docks in Liverpool after years of fighting with the British Navy, he never expects to fall in love. But the moment he glimpses Stella Bennett, all thoughts of returning to his homeland vanish. Determined to be together, the young couple defy Stella’s parents’ wishes. But their plans to marry are soon derailed by the arrival of baby Ginny. Her crop of fiery red hair plants a seed of doubt in her father’s mind, and it’s not long before their world begins to fall apart.

At the heart of Dover's Market Square lies the Castle's Café. A bustling, homely place run by the formidable Nellie Castle and her six children. Since the scandalous birth of her beloved son 10 years before, Marianne, Nellie's eldest daughter, has preferred to stay in the kitchen, hidden away from the town gossips. Overcome with shame, Marianne has never revealed the name of Donny's father - not even to her own mother. But with World War II just around the corner, Marianne's past catches up with her. And suddenly the lives of the Castle family become a lot more complicated.

Hull, 1940. Widowed Edie Copeland has just arrived on Exchange Street, heavily pregnant and with heavy suitcase in tow. She left York after tragedy struck to make a fresh start, but she's a stranger to this street, and her new landlord doesn't hesitate to remind her of this, widow or not. Luckily, the neighbours are a little more welcoming and Edie is soon made to feel at home by the Maguires and the Scuttles. As air raids sound and the war feels closer than ever, the community has to stick together. But Edie is hiding something, and she doesn't know how much longer her secret is safe.

 1936. Polly's guardian angel has to work overtime when her large family is forced to move from the countryside they love into central Liverpool. Money is desperately short, and with her mother working and her father sick, Polly is easily led astray by the handsome Sunny Andersen. But soon war looms, and Sunny joins the navy to train as a signaller. After the horrors of the May Blitz, Polly, too, decides she wants to help her country and goes into the WRNs. She hears that an old pal, Tad Donoghue from the Dublin slums, is now in the Royal Air Force.

August 1940: As the Luftwaffe swarm over Liverpool, Shane Quinn decides to move his family back to the safety of Ireland. But his only child, the beautiful Dana, would rather stay and serve her country than flee to a foreign land. Determined to make it on her own, she joins the WAAF with newfound pals Patty and Lucy. There’s plenty of excitement to be had on a RAF station, even a chance or two at love.

Minnie Maddox cares deeply for mothers and their babies - she makes a living by taking in unwanted children and finding them good adoptive homes - and is delighted for her neighbour when she finally becomes a mother after decades of trying. But when the baby dies of natural causes while under her roof, and knowing her neighbour will be devastated, Minnie swaps it with one of the infants in her care. Now 17, Jewel Nightingale knows nothing of her true origins. But assaulted by her hateful cousin and making the dreadful discovery that she is pregnant, she faces a desperate dilemma.

Jo Melling has arrived in Birch End from Australia, still grieving her father's recent death. She's not intending to stay long, but after tracking down her distant family, Jo becomes more involved in village life than she could ever have imagined - and suddenly in danger too. Jo also finds herself drawn to Nick, a handsome newcomer to the village. Nick had planned to settle in Birch End and start a business, but as he grows closer to Jo, he realises he may have to choose between his dreams and a chance at love. 

Holding the letter in trembling hands, Daisy’s future crumbled before her – the words engraved on her heart forever. The village of Little Creek, the long winter of 1867. The first flakes of snow are falling when Daisy Marshall, secretly engaged to her master's son, finds herself jilted at the altar. Heartbroken, Daisy flees to the small village of Little Creek, nestled on the coast of Essex. There she is warmly welcomed – but the village is poverty-stricken, suffering under a cruel lord of the manor. And when cholera hits, the villagers are truly in dire straits.

Carrie Preston is used to hard work - oldest of 10, with an incompetent mother and a drunken father, she struggles to find food for her siblings. But nothing could have prepared her for the day fate deals her family the cruellest of hands, rendering them more desperate than ever. In a harsh world, small acts of kindness can bring great hope - Eli Beckett's offer of a job at the Dragon pub promises a better future for Carrie, especially as Eli's uncle has offered his nephew some spare land to build a music room.

Jessie Burton is lively and intelligent - she wants more than the respectable life in service her mother has planned for her. Times are changing; railways are being built across the land, bringing new freedom and possibilities. Jessie tastes that freedom when she meets an ambitious young navvy, newly arrived in Yorkshire. The attraction between them is overwhelming. And Jared Wilde is determined to make Jessie his wife. 

England, 1937: After a devastating childhood at Blakely Hall Orphanage, 15-year-old Sandra is released. She finds work as a housemaid, finally able to put her past behind her. But the start of World War Two throws the country into turmoil, and her brother, Alf, is sent away to fight, leaving her completely alone. Germany, 1939: Eleven-year-old Frieda is about to board a ship bound for England with her brother, Kurt. But at the last moment, as Frieda stands on the deck crammed with frightened children, she spots her brother jumping off, back to land.

At 18, Cathie longs for more than life as a settler in the Australian bush. So when her uncle offers her the chance to send her to England, she grabs the opportunity with both hands and runs away from her family. But little does she know that he is using her to get revenge on his sister, Liza. Attacked at the docks in Liverpool, Cathie takes refuge with the man who saved her, a man who has his own troubles. But as she slowly regains her memory and meets her Lancashire relatives, she must confront the legacy of her mother's past.