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Florence, 1415. Banker Cosimo Medici is looked down upon as a jumped up money-changer. Then he marries the noble Contessina de Bardi and becomes the richest man in Europe. But his wealth and power threaten the Florentine elite, who are determined to bring him down. However, the House of Medici cannot be crushed that easily. Sixty years and many violent deaths later, the family are the unquestioned, ruthless rulers of Florence.

When Dr Anna Diaz moves from an urban practice in Cardiff to take on a new job in Bly, a remote village in rural Wales, she finds a community reeling from the unexpected death of their last GP and a startling number of curious rare medical conditions. Among them, a young woman who has become so hairy she looks like an ape, a farmer who is growing new bones, a teacher who hasn't slept for a month, and an old man who has gained enormous amounts of weight and has horns growing out of his head. Deeply unsettled, the locals feel something viral and horrendous is happening to them...